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Fig 1. Male genitalia of Eristalinus. A – E. aeneus, ventral view (1), hypandrium, lateral view (2), apical part of surstylus, ventral view (3); B – E. sepulchralis, ventral view (1), surstylus, lateral view (2), hypandrium, lateral view (3); C – E. taeniops, ventral (1) and lateral view (2); D – E. megacephalus, ventral (1) and lateral view (2). A and B modified from Kanervo (1938). al – aedeagal lobe; bvl – basoventral lobe of surstylus; c – cercus; dpa – dorsal part of the aedeagal lobe; h – hypandrium; s – surstylus; sl – superior lobe of hypandrium. sequencer. The primers used for amplifying and sequencing COI and the D2-3 region of 28S are listed in Table 2. The sequences were assembled and inspected using Sequence NavigatorTM(version 1.01). For this paper we sequenced mainly European species of Eristalinus but some taxa from the Afrotropical region were also sequenced. In the COI gene there was no ambiguity caused by gaps, the D2-3 region of the 28S gene was manually aligned. Parsimony analysis was performed on the combined data matrix using the program Nona vs. 2.0 (Goloboff, 1993), (with commands hold*; hold/20; mult*200) and branch support (Bremer support) was calculated using the same program. Eumerus etnensis van der Goot, 1964 (Eumerini) was used as outgroup. RESULTS DESCRIPTION OF THE MALE GENITALIA

We include here a full description of the male genitalia of E. megacephalus as Shatalkin (1971) provides only a partial drawing. As some pieces of the hypandrium of E. taeniops are not properly detailed in the figures in Kanervo’s paper (1938), we also include a description of the genitalia of this species. Eristalinus taeniops (Wiedemann, 1818) Ventral view (Fig. 1C1)

The inner sides of the surstylus are noticeably sinuous. Superior lobes clearly with sinuous inner and upper sides and with acute and peak shaped extremities. The dorsal part of the aedeagus is regularly convex. Lateral view (Fig. 1C2) The surstylus of Eristalinus taeniops in lateral view has a very prominent and well-developed basoventral lobe. Hypandrium of Eristalinus taeniops evenly broad, with the superior lobe rounded apically and with one conspicuous subventral projection. Aedeagal lobe ventrally incurved with a small ventral hook. Material examined. 5 males, Thermi (Lesbos) Greece coll. 8.IV.2001, leg.: C. Pérez-Bañón & S. Rojo (CEUA); 2 males, San Vicente del Raspeig (Alicante) Spain, coll. VI.1995, leg.: J. Ordóñez (CEUA); 2 males, El Torno (Cáceres) Spain, coll. 25.VIII.1980, leg.: M. A. Marcos-García (CEUA). 1 male, Luxor Egypt (44628. II), Eristalis taeniops Wiedem det.: Becker (FMNH); 1 male, Ishurum ? 1.4.36. F. Brandt, C.1.4. (FMNH); 2 male, 505, 507, Gr. Canaria, Tafira, R. Frey (FMNH).

Eristalinus megacephalus (Rossi, 1794) Ventral view (Fig. 1D1)

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